This question looks old, I faced the same recently.
It seems like after a few months it removes testers once all builds expire from a test flight.
It was fixed after I added a new test flight builds, it took longer than normal before it notify my internal testers approx 2hours but, eventually, it added them back and notify them of the new build version.
No need to delete all users and re invite.
Topic:
App Store Distribution & Marketing
SubTopic:
TestFlight
Tags: